| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| American Century Investment Management, Inc. | 5.9% | 3,610,095 | 3,483,147 | American Century Investment Management, Inc. | /s/ John Pak / Senior Vice President | ||
| American Century Companies, Inc. | 5.9% | 3,610,095 | 3,483,147 | American Century Companies, Inc. | /s/ John Pak / Senior Vice President | ||
| Stowers Institute for Medical Research | 5.9% | 3,610,095 | 3,483,147 | Stowers Institute for Medical Research | /s/ Richard W. Brown / Chairman |