| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| EcoR1 Capital, LLC | 10% | 11,578,447 | 0 | 11,578,447 | /s/ Oleg Nodelman | Manager | |
| Oleg Nodelman | 10% | 11,578,447 | 0 | 11,578,447 | /s/ Oleg Nodelman | Reporting person | |
| EcoR1 Capital Fund Qualified, L.P. | 9.4% | 10,918,474 | 0 | 10,918,474 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC |