| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| EcoR1 Capital, LLC | 9.9% | 5.3M | 0 | 5.3M | /s/ Oleg Nodelman | Manager | 0001587114 |
| Oleg Nodelman | 9.9% | 5.3M | 0 | 5.3M | /s/ Oleg Nodelman | Reporting person | 0001454385 |
| EcoR1 Capital Fund Qualified, L.P. | 9.2% | 4.97M | 0 | 4.97M | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC | 0001598292 |