| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| EcoR1 Capital, LLC | 5.8% | 2,892,000 | 0 | 2,892,000 | /s/ Oleg Nodelman | Manager | |
| Oleg Nodelman | 5.8% | 2,892,000 | 0 | 2,892,000 | /s/ Oleg Nodelman | Reporting person | |
| EcoR1 Capital Fund Qualified, L.P. | 5.4% | 2,709,498 | 0 | 2,709,498 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC |