| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Sarepta Therapeutics, Inc. | 9.5% | 11.9M | 0 | 11.9M | /s/ Ian Estepan | Ian Estepan, Executive Vice President, Chief Financial Officer | 0000873303 |
| Sarepta Therapeutics Investments, Inc. | 9.5% | 11.9M | 0 | 11.9M | /s/ Joe Bratica | Joe Bratica, President |