| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Syncona Portfolio Limited | 11% | 28,821,053 | 0 | 28,821,053 | /s/ Nicholas Moss | Nicholas Moss, Director | |
| Syncona Holdings Limited | 11% | 28,821,053 | 0 | 28,821,053 | /s/ Nicholas Moss | Nicholas Moss, Director | |
| Syncona Investment Management Limited | 11% | 28,821,053 | 0 | 28,821,053 | /s/ Christopher Hollowood | Christopher Hollowood | |
| Syncona Limited | 11% | 28,821,053 | 0 | 28,821,053 | /s/ Robert Hutchinson | Robert Hutchinson | |
| Roel Bulthuis | 11% | 28,821,053 | 0 | 28,821,053 | /s/ Roel Bulthuis | Roel Bulthuis | |
| Christopher Hollowood | 11% | 28,821,053 | 0 | 28,821,053 | /s/ Christopher Hollowood | Christopher Hollowood | |
| Kenneth Galbraith | 11% | 28,821,053 | 0 | 28,821,053 | /s/ Kenneth Galbraith | Kenneth Galbraith |