abrdn plc - Jun 30, 2025 SCHEDULE 13G Report for CODEXIS, INC. Common Stock, par value $0.0001 per share (CDXS)

Filing Manager: Aberdeen Group plc
Reporting Manager: Aberdeen Group plc
Security: CODEXIS, INC. Common Stock, par value $0.0001 per share
Symbol: CDXS
Shares outstanding: 82.9M shares
Disclosed Ownership: 4.2M shares
Ownership: 5.07%
Form type: SCHEDULE 13G
Date filed: 7/30/2025, 11:24 AM
Date of event: Jun 30, 2025
SEC: View on sec.gov

Reporting Managers (2)

Name	Ownership	Shares Owned	sole voting power	shared voting power	Signature	Title	CIK
Aberdeen Group plc	5.07%	4.2M	0	4.2M	Jenny Clarke	Major Shareholding Reporting Analyst
abrdn Inc.	5.07%	4.2M	0	4.2M	Jenny Clarke	Major Shareholding Reporting Analyst