| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 11.3% | 2.9M | 0 | 2.9M | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 8.6% | 2.19M | 0 | 2.19M | Kristin Mariani | Head of North America Compliance, CCO |