| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 12.1% | 3.49M | 0 | 3.49M | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 10.6% | 2.96M | 0 | 2.96M | Kristin Mariani | Head of North America Compliance, CCO |