| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 10% | 170,097 | 0 | 170,097 | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 6.6% | 118,196 | 0 | 118,196 | Kristin Mariani | Head of North America Compliance, CCO |