| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| RA Capital Management, L.P. | 9.9% | 1.61M | 0 | 1.61M | /s/ Peter Kolchinsky | By Peter Kolchinsky, Authorized Signatory | |
| Peter Kolchinsky | 9.9% | 1.61M | 0 | 1.61M | /s/ Peter Kolchinsky | Peter Kolchinsky | |
| Rajeev Shah | 9.9% | 1.61M | 0 | 1.61M | /s/ Rajeev Shah | Rajeev Shah | |
| RA Capital Healthcare Fund, L.P. | 9.7% | 1.61M | 0 | 1.56M | /s/ Peter Kolchinsky | By RA Capital Healthcare Fund GP, LLC, its General Partner, By Peter Kolchinsky, Manager |