| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| EcoR1 Capital, LLC | 6.9% | 6,728,116 | 0 | 6,728,116 | /s/ Oleg Nodelman | Manager | |
| Oleg Nodelman | 6.9% | 6,728,116 | 0 | 6,728,116 | /s/ Oleg Nodelman | Reporting person | |
| EcoR1 Capital Fund Qualified, L.P. | 6.5% | 6,283,426 | 0 | 6,283,426 | /s/ Oleg Nodelman | Manager of the General Partner, EcoR1 Capital, LLC |