| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| Perceptive Advisors LLC | 1.6% | 1.48M | 0 | 1.48M | /s/ Joseph Edelman | Joseph Edelman, Managing Member | |
| Joseph Edelman | 1.6% | 1.48M | 0 | 1.48M | /s/ Joseph Edelman | Joseph Edelman | |
| Perceptive Life Sciences Master Fund, Ltd. | 1.6% | 1.48M | 0 | 1.48M | /s/ Joseph Edelman | Joseph Edelman, Managing Member |