| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 19% | 1,172,368 | 0 | 1,172,368 | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 16% | 1,006,928 | 0 | 1,006,928 | Kristin Mariani | Head of North America Compliance, CCO |