| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 13% | 4,339,254 | 0 | 4,339,254 | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 7.6% | 2,503,168 | 0 | 2,503,168 | Kristin Mariani | Head of North America Compliance, CCO |