| Name | Ownership | Shares Owned | sole voting power | shared voting power | Signature | Title | CIK |
|---|---|---|---|---|---|---|---|
| JANUS HENDERSON GROUP PLC | 14% | 4,557,920 | 0 | 4,557,920 | Kristin Mariani | Head of North America Compliance, CCO | |
| Janus Henderson Biotech Innovation Master Fund Ltd | 8.1% | 2,683,874 | 0 | 2,683,874 | Kristin Mariani | Head of North America Compliance, CCO |